They are much more "regular shoe" in feel than KSOs or Treks. They feel like a very, very flexible running or court shoe. Very comfortable, once I got the lacing to the right tightness.
I have very wide feet and a longer second toe than my big toe. So I always have some compromise to make when I get VFFs, because the toes don't really match my toes. I am a 45 in KSOs and Treks, but I had to go to a 46 in the Speeds because they don't seem to have as much leeway in terms of width as the Treks do (the sidewalls are thicker, so I think they are less stretchy). But the 46s feel good. I have the same problem with these as I have with all my VFFs, which is that the end of my little toe is so much shorter than my second toe that it only fills about half the toe pocket and slips out easily if I scrunch my foot. But I'm used to that and it doesn't really affect the mechanics of the shoe, nor is it an issue with this particular model over others.
I am really liking the laces over the traditional VFF strap system. The shoe feels much more securely fastened to my foot, without sacrificing any flexibility in the sole or toes. Less side-to-side slippage under load and noticeably better stability front-back too.
I ran and walked in them a little today, both on pavement and dirt, and up and down some steep dirt inclines. They didn't shift on me at all, and I had pretty decent ground feel. Not as good as a pair of Classics or KSOs (the sole is a bit thicker), but good.
This is the first pair I have had with the "anatomical pod" Bikila-type outsole and I am sold on it. They're more flexible than my Treks but feel almost as good at protecting from pokey things. They also feel really light. Plus I like the style! I'll definitely be throwing these on with jeans for going out to dinner or cruising around town. My only caveat is that I don't want to mess them up with too much crashing through bushes and dirt, because they look so nice with their shiny black-and-white motif. So I'll probably keep my Treks as my go-to shoe for more outdoorsy stuff, and use the Speeds for city life.
Overall: loving them so far. Probably my favorite pair of VFFs overall. As far as whether they're better than KSOs? If you're mostly cruising around in the civilized world, probably, if only for aesthetics. For serious rough-terrain hiking I'd still prefer my Treks for the little bit of additional armor and the durability of the leather. KSOs I really like as a lighter option for hiking too. Definitely try them out and see how they feel. The only VFFs I have had that I didn't use the hell out of were my Classics, because I really need the straps/laces to get the fit right with my funny feet.
EDIT: Timothy, I think you are my only reader!